Coenobita possess compound eyes comprised of faceted lenses which are especially adept at picking up fine movements. The eyes are located on movable stalks and this is one of the primary ways we determine the species of hermit crab we are looking at.
Reconstructed brains of Fuxianhuia protensa and land hermit crab Coenobita clypeatus showing homologies with Malacostraca. They share 3 nested neuropils in each eye stalk (arrowed 1-3), A1n (antennal nerve), A2n (second pair of nerve roots), op t (optic tract).
Hermit Crab Antenna Eyes Diagram by Storm Martin 2012
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Overview of the anatomy of a land hermit crab (Coenobita)
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Sean Carroll
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